Pebbled Pixie Cup

(Cladonia pyxidata)

Conservation Status

Pebbled Pixie Cup
Dan W. Andree
IUCN Red List

not listed

 
NatureServe

NNR - Unranked

SNR - Unranked

 
Minnesota

not listed

 
     

Description

Pebbled Pixie Cup is a common, cosmopolitan lichen. It occurs worldwide on every continent including Antarctica. In the United States it is common in the east and in the west, but it is mostly absent from the deep south, the desert southwest, the Great Plains, and the Great Basin. It grows under full sun or partial shade, on soil, especially on acidic soil and thin soil over rocks, and it rarely grows on wood.

Pebbled Pixie Cup produces two types of vegetative growth (thallus). The primary thallus is a leaf-like scale (squamule) that lies on the soil. The squamules are numerous but often small and inconspicuous, 116 to 316 (2 to 5 mm) long, 132 to (1 to 3 mm) wide, and very thin, less than 164 (0.4 mm) thick. The margins may be irregularly lobed, toothed, cut (incised), or smooth (entire). The lobes are mostly ascending, and they never form a rosette-like crust. The squamules are persistent, remaining as the secondary thallus matures.

The secondary thallus is a vertical, unbranched, greenish gray to brown stalk (podetium) that is to ¾ (3 to 20 mm) tall and rises from the primary thallus. The podetium broadens gradually at the tip into a 516 to ½ (8 to 12 mm) in diameter cup. A key characteristic identifying this species is the podetium widening gradually from the base to the cup without a distinct, narrow stalk.

The cup is undivided and the margins are unlobed. The layer of protective tissue (cortex) may be complete up to the cup, just at the base, or entirely missing. When present, the cortex is warty, and it breaks up into granules or gives rise to tiny, asexual, reproductive structures. When these are formed in the cup, they may appear like appressed squamules, a feature characteristic even in young specimens.

Egg-shaped, reproductive, flask-like structures (pycnidia) are very common on the margins. Disk-shaped reproductive structures (apothecia) on the margins of the cup are also common. They are up to 516 (8 mm) wide, on stalks about (3 mm) long, and reddish brown to a darker color.

Pebbled Pixie Cup contains fumarprotocetraric acid, which causes a red reaction during standard lichen spot testing (the P+ red test).

Similar Species

 

Ecology

Substrate

Ground

Growth Form

Fruticose

Habitat

 

Hosts

 

Distribution

Distribution Map

 

Sources

24, 26, 29, 30, 77, 81. iNaturalist.org (30) temporarily unavailable

Biodiversity occurrence data published by: Minnesota Biodiversity Atlas (accessed through the Minnesota Biodiversity Atlas Portal, bellatlas.umn.edu, 10/30/2025).

10/30/2025    
     

Occurrence

Common and widespread

Taxonomy

Kingdom

Fungi (fungi)

Subkingdom

Dikarya

Phylum

Ascomycota (sac fungi)

Subphylum

Pezizomycotina (sac fungi amd lichens)

Class

Lecanoromycetes (common lichens)

Subclass

Lecanoromycetidae (shield lichens, sunburst lichens, rosette lichens, and allies)

Order

Lecanorales (shield lichens, rim lichens, and allies)

Suborder

Lecanorineae

Family

Cladoniaceae (spindles and structured lichens)

Genus

Cladonia (pixie cup and reindeer lichens)

Mycobiont

Cladonia fimbriata

Photobiont

Subordinate Taxa

 

Synonyms

Baeomyces neglectus

Baeomyces pyxidatus

Baeomyces pyxidatus var. staphyleus

Capitularia neglecta

Cenomyce neglecta

Cenomyce pyxidata

Cenomyce pyxidata var. syntheta

Cladonia chlorophaea ssp. costata

Cladonia chlorophaea var. costata

Cladonia communis

Cladonia conchata

Cladonia floccida

Cladonia neglecta

Cladonia neglecta ssp. scyphosa

Cladonia neglecta var. scyphosa

Cladonia polymorpha var. pyxidata

Cladonia polymorpha var. tuberculata

Cladonia pyxidata ssp. lophyra

Cladonia pyxidata ssp. neglecta

Cladonia pyxidata ssp. simplex

Cladonia pyxidata var. epiphylla

Cladonia pyxidata var. floccida

Cladonia pyxidata var. lophyra

Cladonia pyxidata var. neglecta

Cladonia pyxidata var. pachyphyllina

Cladonia pyxidata var. polyscypha

Cladonia pyxidata var. pyxidata

Lichen polymorphus var. pyxidatus

Lichen pyxidatus

Lichen pyxidatus var. tuberculatus

Lichen tuberculatus

Patellaria neglecta

Patellaria pyxidata

Scyphophorus pyxidatus

Common Names

Pebbled Pixie Cup

Pebbled Pixie-Cup

Pebbled Pixie-Cup Lichen

 

 

 

 

 

Glossary

Apothecium

An open, disk-shaped or cup-shaped, reproductive structure, with spore sacs on the upper surface, that produces spores for the fungal partner of a lichen. Plural: apothecia.

 

Fruticose

Shrubby: referring to the growh form of lichens that may be tufted, draped, or stalked.

 

Podetium

The hollow stalk of the fruiting body of lichens in the genus Cladonia. Plural: podetia.

 

Pycnidium

A small, spherical, flask-shaped, or inversely pear-shaped, asexual reproductive structure produced by fungi in which conidia develop. Plural: pycnidia.

 

Soredium

An asexual reproductive structure of a lichen in the form of a tiny dull granule on the thallus surface that can be easily brushed off. It consists of a cluster of algal cells (the photobiont) wrapped in fungal filaments (the mycobiont), but without an outer layer of protective tissue (cortex). Plural: soredia.

 

Squamules

On lichens: small, flat, often overlapping, leaf-like scales without a lower cortex. Adjective: squamulose

 

Thallus

The vegetative body of a lichen composed of both the alga and the fungus.

 

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Nancy Falkum

Pebbled Pixie Cup

Dan W. Andree

Pebbled Pixie Cup

Another type of Lichen...

This interesting lichen caught my eye recently. I had seen it in spots but usually smaller growths. This one had nice light green color and like a bunch of little cup like protrusions sticking upward. A more common and abundant in areas compared to the reindeer lichens. Really interesting and nice color etc.

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Other Videos

Pebbled Pixie-Cup Lichen!
Pryce Durnye

About

May 13, 2021

In which, I am lichen me some nature! In the video, I introduce the Pebbled Pixie-Cup Lichen. The clue is in the name. The lichens resemble looking at a cup or chalice. They are another example of a fruticose lichen that has podetia (upright structure).

Also, take a moment to observe the beloved waterfalls :D! There are honestly not many sources that specifically focus on this species but here's what I do possess.

 

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Visitor Sightings

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Dan W. Andree
10/24/25

Pebbled Pixie Cup

Location: Norman Co. Mn.

Nancy Falkum
10/23/25

Pebbled Pixie Cup

Location: Superior National Forest, Ram Lake Trail

MinnesotaSeasons.com Sightings

 

 

 

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