water speedwell

(Veronica catenata)

               
Conservation Status

IUCN Red List

LC - Least Concern

water speedwell

 

NatureServe

NNR - Unranked

SNR - Unranked

Minnesota

not listed

Wetland
Indicator
Status

Great Plains

OBL - Obligate wetland

Midwest

OBL - Obligate wetland

Northcentral & Northeast

OBL - Obligate wetland

Nativity

Native

Occurrence

Common and widespread

Habitat

Wet. Springs, slow streams, edges of ponds, and ditches. Full to partial sun. Muddy or sandy and gravelly soil.

Flowering

May to September

 
Flower Color

White to pink or pale bluish with purple streaks

Height

8 to 40

 
 
Identification

Water speedwell is a large semi-aquatic plant. It occurs across the United States and southern Canada. It is common in Minnesota. It grows in full sun, in muddy or sandy and gravelly soil, in springs, slow streams, edges of ponds, and ditches.

Due to the uncertain taxonomy of this species, its distribution is uncertain. Some reputable sources believe this is the European species blue water speedwell (Veronica anagallis-aquatica). They contend that it is not native but was introduced into North America and is now widely naturalized. Other equally reputable sources insist that this is a separate species native to North America. They contend that the two species are easily separable by visible characteristics. Most sources list only one of the two species in North America. A few show both species occurring in the United States. In either case, there is intergradation of the characteristics and separation of the two species in the field is not as clear as the plant identification dichotomous keys would make it seem. Making matters worse, the two plants readily hybridize.

Water speedwell is an erect, perennial forb that rises on multiple stems from a thin, underground, horizontal stem (rhizome) and fibrous roots. It can be 8 to 40 tall but is usually no more than 24 in height. It is often emergent, with the base of the plant under water. Sometimes it is completely submerged.

The stems are usually erect or curve up strongly from the base (ascending). In areas with flooding they may be nearly horizontal (spreading). They are branched and usually completely hairless. Sometimes they are sparsely to moderately covered with minute, gland-tipped hairs just in the inflorescence. Roots develop at the lowest nodes.

The leaves are opposite, ¾ to 3 long, ¼ to 1½ wide, mostly lance-shaped (widest below the middle), sometimes inversely lance-shaped (widest above the middle) or oblong (widest and with straight sides along the middle). Most are 3 to 5 times as long as wide, though some may be up to 8 times as long as wide, especially just below the inflorescence. Most are stalkless but the lower leaves sometimes have short leaf stalks. The leaf blades are heart-shaped at the base and are mostly sharply pointed, sometimes bluntly pointed at the tip. The stalkless leaves more or less wrap partially around (clasp) the stem at the base. The upper surface is hairless and minutely pitted, though this requires a hand lens to see. The lower surface is mostly hairless but may have inconspicuous gland-tipped hairs near the base of the midrib. The margins may be untoothed or have minute, widely-spaced teeth, especially near the tip.

The inflorescence is a solitary unbranched cluster (raceme) of 15 to 25 flowers rising from each of the upper leaf axils. The raceme is up to about 2¾ (7 cm) long and is on a hairless stalk (peduncle) that is up to 19 16 (4 cm) long. Each flower is on a spreading to ascending, to 3 16 (2.5 to 5 mm) long stalk (pedicel). When in fruit, the pedicel lengthens to ¼ (7 mm) and spreads at a right angle to the axis of the raceme or occasionally slightly curves upward. At the base of each pedicel there is a single, small, modified leaf (bract). The bract is narrow, 1 16 to (1.5 to 4 mm) long, 1 16 wide, and straight-sided (linear) or lance-shaped.

The flowers are to ¼ (3 to 6 mm) in diameter. There are 4 outer floral leaves (sepals), 4 petals, 2 stamens, and 1 style. The sepals are green and (3.0 to 3.5 mm) long. They are hairless or may have minute, gland-tipped hairs toward the base. They are fused at the base into a short, 1 32 (1 mm) long calyx tube, then separated into 4 broadly lance-shaped to egg-shaped lobes. The petals (corolla) are fused at the base into a short tube then separated into 4 lobes that are much longer than the tube. The throat is green inside and there is a narrow white ring where the lobes separate. The corolla lobes are broadly egg-shaped, (3 mm) long and wide, and slightly curved upward. The upper and lateral lobes are equal in size, the lower lobe is smaller. All of the lobes are white to pink or pale bluish with darker purple veins. On the upper lobe and the upper half of the lateral lobes the veins are even darker and widen at the base. The stamens are (2.5 to 3.0 mm) long and project well beyond the petals. The stalks of the stamens (filaments) are white, becoming tinged with purple toward the tip. The style is (3 mm) long and also projects well beyond the petals. It is also white at the base and purplish tinged toward the tip.

The fruit is a slightly flattened, (3 to 4 mm) long and wide capsule with numerous tiny seeds. The capsule is broadly triangular or slightly heart-shaped with just a shallow notch at the tip. The surfaces are hairless. The margin may be hairless or have a few gland-tipped hairs.

 
Similar
Species

 

 
Distribution Distribution Map  

Sources: 2, 3, 4, 5, 7, 22, 28, 29, 30.

 
Comments

Taxonomy
Some sources, including BONAP2 and USDA PLANTS3, consider this plant the same as the European species blue water speedwell (Veronica anagallis-aquatica). Most sources list only one of the two species in North America. At least three sources, including Gleason & Cronquist41, GBIF29 and iNaturalist, show both species occurring in the United States. In plant identification dichotomous keys the two species are easily separable by visible characteristics. In the field, however, there is intergradation of the characteristics and separation of the two species is not as clear. Making matters worse, the two plants readily hybridize.

 
Taxonomy

Family:

Plantaginaceae (plantain)

 

Tribe:

Veroniceae

 

Genus:

Veronica

 

Subgenus:

Beccabunga

 
Subordinate Taxa

Some authors recognize the variety var. glandulosa. These plants have gland-tipped hairs on the stem, whereas the nominate variety (var. catenata) stems are hairless. The fruit is sometimes slightly narrower. Most authors consider the differences too slight to warrant separation, and recognize no varieties.

 
Synonyms

Veronica catenata var. glandulosa

Veronica comosa var. glaberrima

Veronica comosa var. glandulosa

Veronica connata

 

Veronica connata ssp. glaberrima

Veronica connata var. glaberrima

Veronica connata var. typica

 
Common
Names

chain speedwell

Pennell’s speedwell

pink water-speedwell

 

sessile water-speedwell

water speedwell

 

 

 

 

 

 

 

 

Glossary

Ascending

Curving upward from the base.

 

Axil

The upper angle where a branch, stem, leaf stalk, or vein diverges.

 

Bract

Modified leaf at the base of a flower stalk or flower cluster.

 

Clasping

Describing a leaf that wholly or partly surrounds the stem but does not fuse at the base.

 

Filament

On plants: The thread-like stalk of a stamen which supports the anther. On Lepidoptera: One of a pair of long, thin, fleshy extensions extending from the thorax, and sometimes also from the abdomen, of a caterpillar.

 

Glandular hairs

Hairs spread over aerial vegetation that secrete essential oils. The oils act to protect against herbivores and pathogens or, when on a flower part, attract pollinators. The hairs have a sticky or oily feel.

 

Linear

Long, straight, and narrow, with more or less parallel sides, like a blade of grass.

 

Node

The small swelling of the stem from which one or more leaves, branches, or buds originate.

 

Pedicel

In plants: the stalk of a single flower in a cluster of flowers. In Hymenoptera and Araneae: the narrow stalk connecting the thorax to the abdomen.

 

Peduncle

In angiosperms, the stalk of a single flower or a flower cluster; in club mosses, the stalk of a strobilus or a group of strobili.

 

Raceme

An unbranched, elongated inflorescence with stalked flowers. The flowers mature from the bottom up.

 

Rhizome

A horizontal, usually underground stem. It serves as a reproductive structure, producing roots below and shoots above at the nodes.

 

Sepal

An outer floral leaf, usually green but sometimes colored, at the base of a flower.

 

Spreading

Extending nearly horizontal.

       
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Plant

  water speedwell   water speedwell
       

Inflorescence

  water speedwell   water speedwell
       
  water speedwell    
       

Flowers

  water speedwell   water speedwell
       

Leaves

  water speedwell   water speedwell
       

 

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  Veronica anagallis aquatica is a species of flowering plant in the family Plantaginaceae
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About

May 21, 2019

I go through the forests, mountains, hills, fields, and waters to understand the living world and to create a living mind.

Veronica anagallis-aquatica is a species of flowering plant in the family Plantaginaceae known by the common names water speedwell, blue water-speedwell, brook pimpernel, and sessile water-speedwell, Veronica catenata.

   
       

 

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Created: 12/20/2019

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