Mealy Pixie Cup

(Cladonia chlorophaea)

Information

Mealy Pixie Cup - Species Profile

Mealy Pixie Cup - Featured photo
Dan W. Andree

Conservation Status

IUCN Red List

not listed

NatureServe

NNR - Unranked
SNR - Unranked

Minnesota

not listed

Description

Mealy Pixie Cup is a common, cosmopolitan lichen. It occurs worldwide on every continent except Antarctica. In the United States it is common in the east and in the west, but it is mostly absent from the deep south, the desert southwest, the Great Plains, and the Great Basin. It grows under full sun or partial shade, on humus and mineral soil, on rotten wood, and on tree trunks.

Mealy Pixie Cup produces two types of vegetative growth (thallus). The primary thallus consists of numerous leaf-like scales (squamules) that often form a dense, carpet-like mat on the substrate. The squamules are to ¼ (3 to 6 mm) long and 116 to (2 to 4 mm) wide, and they are ascending or raised at the margins.The margins have irregular, rounded lobes or teeth. The upper side is grayish green and the underside is white, but the color darkens toward the base. The squamules are persistent, remaining as the secondary thallus matures.

The secondary thallus is a vertical, unbranched, stout or more or less slender stalk (podetium) rising from the primary thallus. It is grayish green to brownish-green and 116 to (2 to 15 mm) tall, and does not darken or turn black at the base (non-melanotic). At the tip of the stalk there is a 116 to ¼ (2 to 6 mm) wide cup that gradually tapers to the stalk. The cup is undivided and the margin is unlobed, though it may appear lobed when fruiting structures are present. The layer of protective tissue (cortex) is typically absent or is present just at the very base. As the lichen develops, the cortex breaks up and gives rise to fine, asexual reproductive structures (soredia). These soredia are mealy in texture and cover the outer surface and the interior of the cup.

Egg-shaped to top-shaped (turbinate) reproductive, flask-like structures (pycnidia) are very common on the margin. Disk-shaped, 116 to (2 to 4 mm) in diameter, brown, reproductive structures (apothecia) rising from the margin are uncommon.

Mealy Pixie Cup contains fumarprotocetraric acid, which causes a red reaction during standard lichen spot testing (the P+ red test).

Similar Species

Pebbled Pixie Cup (Cladonia pyxidata) is covered in coarser, pebble-like or scale-like granules. However, identification is not always definitive. Juvenile specimens that haven't fully developed soredia, or very old (senescent) specimens where the surface has eroded, can look identical. Because these two species exist on a morphological gradient, some lichenologists argue they should be treated as a single variable species. Both species are often found growing right next to each other.

Trumpet Lichen (Cladonia fimbriata) soredia are significantly finer, resembling a coating of white flour. The cups are typically taller, narrower, and more “trumpet-shaped.”

Ecology

Substrate

Ground

Growth Form

Fruticose

Habitat

 

Hosts

 

Distribution

Map
3/7/2026

Sources

7, 24, 30, 77, 83.

Biodiversity occurrence data published by: Minnesota Biodiversity Atlas (accessed through the Minnesota Biodiversity Atlas Portal, bellatlas.umn.edu. Accessed 3/7/2026).

The Consortium of North American Lichen Herbaria (CNALH) https://lichenportal.org/cnalh/index.php. Accessed 3/7/2026.

Mycology Collections Portal (MyCoPortal) https://www.mycoportal.org/portal/collections/index.php). Accessed 3/7/2026.

Cladonia chlorophaea (Flörke ex Sommerf.) Spreng. in GBIF Secretariat (2023). GBIF Backbone Taxonomy. Checklist dataset https://doi.org/10.15468/39omei accessed via GBIF.org. Accessed 3/7/2026.

Fink, B. (1910). The Lichens of Minnesota. United States: U.S. Government Printing Office.

Occurrence

Common

Taxonomy

Kingdom

Fungi (Fungi)

Subkingdom

Dikarya

Phylum

Ascomycota (Sac Fungi)

Subphylum

Pezizomycotina

Class

Lecanoromycetes (Common Lichens)

Subclass

Lecanoromycetidae (Shield Lichens, Sunburst Lichens, Rosette Lichens, and Allies)

Order

Lecanorales (Shield Lichens, Rim Lichens, and Allies)

Family

Cladoniaceae (Spindles and Structured Lichens)

Genus

Cladonia (Pixie Cup and Reindeer Lichens)

Mycobiont

Cladonia chlorophaea

Photobiont

green algae other than Trentepohlia

Species

Cladonia chlorophaea was long treated as a subspecies or variety of the Pebbled Pixie Cup (Cladonia pyxidata). Recent phylogenetic studies and taxonomic revisions have seen it raised to full species status.

Species Complex

Modern lichenology treats C. chlorophaea as a species complex rather than a single, uniform species. It belongs to a group of “cryptic species”—taxa that are morphologically identical but chemically and genetically distinct.

In Minnesota, a specimen identified in the field as C. chlorophaea may actually be one of several sibling species, most notably Gray’s Pixie Cup (C. grayi). These cannot be reliably separated by sight; they require Thin-Layer Chromatography (TLC) to detect specific secondary metabolites (acids). Because these species are indistinguishable to the naked eye, many older records and field observations group them under the broad “Mealy Pixie Cup” (C. chlorophaea) umbrella.

Subordinate Taxa

 

Synonyms

Cenomyce chlorophaea

Cladonia chlorophaea

Cladonia chlorophaea var. chlorophaea

Cladonia chlorophaea var. pachyphyllina

Cladonia pyxidata subsp. chlorophaea

Cladonia pyxidata var. chlorophaea

Patellaria fimbriata var. chlorophaea

Common Names

Mealy Pixie Cup

Mealy Pixie Cup Lichen

Mealy Pixie-cup

Photos

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Dan W. Andree

Mealy Pixie Cup XX

Another Pixie Cup Lichen...

Had a little larger cups than most I seen.

Minnesota Seasons Photos

Slideshows

Slideshows

Cladonia chlorophaea - fungi kingdom
Fungi Kingdom

About

Jan 23, 2015

Cladonia chlorophaea - fungi kingdom

Videos

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Other Videos

Pixie Cup or Goblet Lichen (Cladonia chlorophaea aggregate)
Roger Griffith

About

Nov 24, 2014

One of the Pixie Cup or Goblet lichens. A member of the Cladonia group, probably either C.chlorophaea or pyxidata. Growing on rocks at South Grange Farm, Dunlop, East Ayrshire, Scotalnd.

One of most abundant lichens in the Ouray, Colorado area. Cladonia chlorophaea
Mike's thoughts on plants.

About

Aug 16, 2021

Cladonia chlorophaea is wide spread locally, with some morphological plasticity. A forest floor, on humus mostly but also on shaded rocks.

Fijn bekermos - Cladonia chlorophaea
Het Wildepad

About

Feb 1, 2023

The fine cup moss (Cladonia chlorophaea) is a lichen from the Cladoniaceae family. It grows on dead trees, tree stumps, and branches, on the ground, on stone, and on wood.

Sightings

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Dan W. Andree
10/31/25

Mealy Pixie Cup

Location: Norman Co. Mn.

Minnesota Seasons Sightings