freeloader flies

(Family Milichiidae)

Overview

Milichiidae is the family of small to very small flies known as freeloader flies or jackal flies. They occur worldwide on every continent except Antarctica.

There are currently 360 described Milichiidae species in 19 genera in 3 subfamilies worldwide, 36 species in 12 genera in 3 subfamilies in North America north of Mexico, and at least 11 species in 9 genera in 3 subfamilies in Minnesota.

 
freeloader flies (Family Milichiidae) and northern crab spider on great blanketflower
Photo by Dan W. Andree
 

Milichiidae are found in a diverse variety of habitats, including in forests, forest edges, and forest canopies, in open areas and caves, and in stables and houses.

Milichiidae are undoubtedly much more common in Minnesota than the map on this page indicates. Due to their small size and lack of economic importance, they are often overlooked or ignored when found in mass traps. They are uncommon in prepared collections.

Some Milichiidae adults are inquilines, living in the nests of ants or birds, or in the roosts of bats. They scavenge near the mouthparts and anal openings of their host. Some Milichiidae adults are kleptoparasitic, feeding on the prey of spiders or predaceous insects. They wait near the predator, and when a prey is captured, they feed on the body fluids of the prey. This is the source of the common name freeloader flies. A freeloader is defined as “someone who takes advantage of the generosity of others” or “one who depends on another for support without reciprocating”. 

Milichiidae larvae are very small. Most species are saprophagous, feeding on rotting vegetable matter, decaying wood, or decaying bark. Some species are coprophagous, feeding on dung or excrement.

Description

Milichiidae adults are small to very small. They can be 132 to ¼ (1 to 7 mm) in length, but most are no more than (3 mm) in length. Most are shiny and black, some are brown, orange, or yellow, and some appear silvery due to a covering of minute, silvery hairs on the upper side of the abdomen of the male.

On the top of the head, the two postvertical bristles either converge or are parallel. On the face, at least one pair of the lower orbital bristles are bent inward. When viewed from the side, the face is not deeply hollowed out between the antennae and the mouthparts. The antennae have three segments. The second segment is not grooved. The third segment is sometimes very large, and there is a bristle (arista) on the upper surface. On many species, the eyes are red. On most species, the tube-like mouthpart (proboscis) is fairly long, and it is jointed in the middle, knee-like. It is folded below the head when not in use. The sensory hairs around the mouth (vibrissae) are present, but they are weakly developed.

The thorax is not crossed by a continuous groove.

The abdomen is short and wide.

The wings are variable in both shape and color. They have complete first basal, second basal, anal, and discal cells. The anal cell is short, closed, and very small. There is no sub-apical cell. The costa has a break near the humeral cross vein and another near the end of vein R1. There is just one small lobe (calypter) at the base of the wing. If a second lobe is present, it is much reduced.

The fourth leg segment (tibia) on each leg does not have a bristle on the upper side near the tip.

Distribution

Distribution Map

 

Sources

24, 27, 29, 30, 82, 83.

7/29/2024    
Taxonomy

Order

Diptera (flies)

Suborder

Brachycera

Infraorder

Cyclorrhapha

Zoosection

Schizophora

Zoosubsection

Acalyptratae

Superfamily

Carnoidea

   

Subordinate Taxa

Subfamily Madizinae

Genus Desmometopa

Desmometopa flavicoxa

Desmometopa inaurata

Desmometopa m-nigrum

Desmometopa microps

Desmometopa nearctica

Desmometopa nigrifemorata

Desmometopa sordida

Desmometopa tarsalis

Desmometopa varipalpis

Genus Leptometopa

Leptometopa beardsleyi

Leptometopa flaviceps

Leptometopa halteralis

Leptometopa latipes

Leptometopa rufifrons

Genus Madiza

Madiza glabra

 

Subfamily Milichiinae

Genus Eusiphona

Eusiphona cooperi

Eusiphona mira

Genus Milichia

Milichia aethiops

Milichia canariensis

Milichia ludens

Milichia mixta

Milichia patrizii

Milichia pubescens

Milichia speciosa

Genus Milichiella

Milichiella arcuata

Milichiella argyrogaster

Milichiella bimaculata

Milichiella bisignata

Milichiella freyi

Milichiella lacteipennis

Milichiella melaleuca

Genus Pholeomyia

Pholeomyia indecora

 

Subfamily Phyllomyzinae

Genus Aldrichiomyza

Aldrichiomyza agromyzina

Genus Neophyllomyza

Neophyllomyza acyglossa

Genus Paramyia

Paramyia nitens

Paramyia pseudonitens

Genus Phyllomyza

Phyllomyza donisthorpei

Phyllomyza equitans

Phyllomyza longipalpis

Phyllomyza milnei

Phyllomyza pallida

Phyllomyza securicornis

Genus Stomosis

Stomosis vittata

   

Synonyms

Phyllomyzidae

   

Common Names

filth flies

freeloader flies

jackal flies

 

 

 

 

 

 

 

 

 

 

 

 

 

Glossary

Arista

A large bristle on the upper side of the third segment of the antenna of a fly. Plural: aristae.

 

Calypter

On flies: one of two small membranous lobes at the base of the forewing that covers the haltere.

 

Proboscis

The tube-like protruding mouthpart(s) of a sucking insect.

 

Tibia

The fourth segment of an insect leg, after the femur and before the tarsus (foot). The fifth segment of a spider leg or palp.

 

 

 

 

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Dan W. Andree

I became acquainted with the Jackal Flies...

I seen a northern crab spider on a blanket flower it had an insect caught and feeding on it when a few small flies showed up, then more and more. They somewhat circled the crab spider and a couple tried sneaking in and at times the spider would take one of its legs and flick it at the flies.

 

freeloader flies (Family Milichiidae) and northern crab spider on great blanketflower

Anyway I googled small flies surround a crab spider feeding. I seen a lady had a video of a crab spider that caught a honey bee and she called the flies trying to get in on it Jackal Flies. They were doing what I seen some do.

But anyway I filmed it on and off for a good 2-3 minutes since the spider had turned nice colors and the flower etc. But anyway I got some nice footage and when I seen how it is laid out etc. it reminded of a small segment of something Nat. Geo. "somewhat". The photo is a frame grab from 4K video. I just shrunk down smaller.

Crab spiders are neat little spiders and can change color somewhat to their surroundings. Also they are so small and harmless to people. They may take a honey bee or some other insect people wouldn’t want a spider to catch, but they aren’t a threat to any species declining because of them. Crab spiders will also feed on other insects like stink bugs, flies, and numerous others including one I seen in the past had a yellow jacket wasp for lunch. I’m not a fan of yellow jacket wasps.

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Milichiidae
Nigel Jones

Milichiidae

Milichiidae
Rui Andrade

Milichiidae

Milichiidae
ikolai Vladimirov

Milichiidae

 

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Other Videos

Milichiidae gen. sp. cleptoparasitism
プッカ (Pucca)

About

Oct 27, 2019

Xysticus and Apis and Milichiidae gen. sp.

"Meet the Freeloader Flies: The Tiny but Mighty Insects with a Sneaky Side"
mohamed moumaris

About

Apr 18, 2023

Milichiidae is a family of small, non-biting flies commonly known as the freeloader flies or jackal flies. They are found worldwide, with over 230 species described, and are typically 1-5mm in length. Milichiidae are often found in association with other insects, especially bees and wasps, and are known to feed on pollen, nectar, and other materials found in the nests of these insects. Some species are also known to be kleptoparasites, stealing food and other resources from other insects.

The body of Milichiidae is typically elongated, with a distinct head and thorax, and long, slender legs. The wings are usually clear or lightly tinted, with distinct veins and a characteristic pattern of dark spots or stripes. The antennae are also long and slender, and may be bent or curved.

Milichiidae are generally considered to be harmless to humans, and are not known to transmit any diseases or cause any significant damage to crops or other agricultural products. They are, however, important components of many ecosystems, playing important roles in pollination, decomposition, and other ecological processes.

 

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Visitor Sightings
 

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Dan W. Andree
Summer 2024

Location: Norman Co. Mn.

I seen a northern crab spider on a blanket flower it had an insect caught and feeding on it when a few small flies showed up, then more and more. They somewhat circled the crab spider and a couple tried sneaking in and at times the spider would take one of its legs and flick it at the flies.

freeloader flies (Family Milichiidae) and northern crab spider on great blanketflower
 

Anyway I googled small flies surround a crab spider feeding. I seen a lady had a video of a crab spider that caught a honey bee and she called the flies trying to get in on it Jackal Flies. They were doing what I seen some do.

But anyway I filmed it on and off for a good 2-3 minutes since the spider had turned nice colors and the flower etc. But anyway I got some nice footage and when I seen how it is laid out etc. it reminded of a small segment of something Nat. Geo. "somewhat". The photo is a frame grab from 4K video. I just shrunk down smaller.

Crab spiders are neat little spiders and can change color somewhat to their surroundings. Also they are so small and harmless to people. They may take a honey bee or some other insect people wouldn’t want a spider to catch, but they aren’t a threat to any species declining because of them. Crab spiders will also feed on other insects like stink bugs, flies, and numerous others including one I seen in the past had a yellow jacket wasp for lunch. I’m not a fan of yellow jacket wasps.

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Created: 7/30/2024

Last Updated:

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